Which of the following leaf modifications is beneficial in a low-light environment, e.g., the understory of a rainforest? (More than one choice may apply.)

(A) Iridescent and reflective.
(B) Undersides are red from high levels of anthocyanin pigments.
(C) Covered in hairs.
(D) Have a LARGE surface area.
(E) Have very SMALL surface area.
(F) Sharp (nonphotosynthetic) spines.